Stauromedusae of the genus *Manania* (=*Thaumatoscyphus*) (Cnidaria, Scyphozoa) in the northeast Pacific, including descriptions of new species *Manania gwilliami* and *Manania handi*

Abstract

*Manania gwilliami* n.sp. and *Manania handi* n.sp., stauromedusae beloning to the familyi Depastridae, are described from the west coast of North America. They are differentiated from the one species of the genus previously known from the area, *M. distincta* (Kishinouye, 1910) by shape, color, habitat, and geographic range. Specimens of *M. gwilliami* are generally red and those of *M. handi* are green; both lack the dark brown herringbone patterning of *M. distincta* medusae. *Manania gwilliami* is widely distributed from Mexico to Canada in exposed shallow-water coastal habitats, whereas *M. handi* has been found only off the San Juan Islands and Vancouver Island in protected shallow water. The occurrence of *M. distincta* in the northeast Pacific has been sporadic. The name *Manania* is a senior synonym of *Thaumatoscyphus*.