Revision of sea anemone genus *Epiactis* (Coelenterata: Actiniaria) on the Pacific coast of North America, with descriptions of two new brooding species

Abstract

Two new species of the actiniid genus *Epiactis*, *Epiactis lisbethae* and *Epiactis fernaldi*, are described from the intertidal zone of the San Juan Islands, Washington, U.S.A. The sea anemone genus *Cnidopus* is synonymized with *Epiactis*, and its type species, *C. ritteri*, is restored to the genus in which it was originally described. Thus, four species of *Epiactis* are known on the Pacific coast of North America: external brooders *E. prolifera* (type species of the genus) and *E. lisbethae*, which differ in sexuality and brooding periodicity; and internal brooders *E. ritteri* and *E. fernaldi*, which differ in cnidae, sexuality, and histology. These bring the number of nominal species in the genus *Epiactis* to 19.