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A Revision of the Genus Elmas Blackwelder, 1952 (Coleoptera: Staphylinidae: Staphylininae: Xanthopygina), with a Preliminary Reconstructed Phylogeny of the Species1
Ashe, James S. Chatzimanolis, Stylianos
Ashe, James S.
Chatzimanolis, Stylianos
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Abstract
The staphylinid genus Elmas Blackwelder 1952 (type.species: Selma modesta Sharp 1876
from Chontales, Nicaragua) is revised for the first time. The two previously described species, E. modesta
(Sharp) from Nicaragua and E. strigella (Bernhauer) from Brazil, are redescribed. Fifteen species are
described as new: Elmas brooksi from Ecuador; Elmas costaricensis from Costa Rica; Elmas elassos from
Ecuador; Elmas esmeraldas from Ecuador; Elmas falini from Suriname; Elmas gigas from Peru; Elmas
guianas from-French Guiana; Elmas hanleyi from Costa Rica; Elmas hibbsi from Ecuador; Elmas lambas
from Brazil; Elmas lescheni from Peru; Elmas panamaensis from Panama; Elmas patillas from Costa Rica;
Elmas spinosus from Bolivia; and Elmas windsori from Panama. A key and illustrations of structural
features and aedeagi are provided for identification of the known species. The phylogenetic relationships
of the species of Elmas species are only weakly resolved by the available dataset. Elmas is strongly
supported to be a monophyletic lineage, and E. strigella is the most basal species followed by E. lambos
and E. guianas respectively. E. spinosus + E. falini + E. gigas are strongly supported to be a monophyletic
. group; E. hfbbsi is weakly supported to be the sister group to these three species. The lineage (E. windsori
· + E. costaricensis) + (E. panamaensis + E. patillas) is weakly supported in all trees. The lineage E. elassos +
E. hanleyi + E. esmeraldas is also weakly supported, and the successive approximation analysis hypothesizes
that E. modesta is also a member of this lineage.
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Date
2003-04-29
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Natural History Museum, University of Kansas
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Keywords
Staphylinidae, Staphylininae, Xanthopygina, Elmas, revision, key, new species, phylogeny